GGSB Empirical Track Coursework


Suggested “tracks” for students interested in concentrating in the Empirical track have been developed by the CSAC. (Model Systems, Population Genetics, Human Genetics, Developmental Genetics, and Genomics & Systems Biology) have been developed by the CSAC. A summary of the five suggested tracks is given in Appendix A.  Students are required to consult with their assigned mentor prior to registration each quarter.

A total of four graded electives must be taken, one of which may be a reading course.  The electives can be selected according to the student’s interests and the availability of courses.

Four  Required Courses:

Genetic Analysis of Model Organisms  AND  Genomics and Systems Biology   

Plus One of the Following Two Courses:

Molecular Biology I OR  Molecular Biology II

Plus One  of the Following Four Courses:

Fundamentals of Molecular Evolution OR  Principles of Population Genetics I OR Evolutionary Genomics OR Human Variation & Disease

Empirical Track Course Electives [4 courses]

  • Must take 4 courses (see list of approved electives).
  • Students may petition the CSAC for approval of courses not listed in this handbook as “approved”.
  • At least 3 of the 4 electives are to be taken before the Preliminary Exam.
  • All 4 electives should be taken before the Qualifying Exam.
  • One of the 4 elective courses may be taken pass/fail subject to CSAC approval.
  • One of the electives may be a graded reading course (see guidelines for reading courses).


Click below for a list of GGSB course descriptions, elective and student tracks: